As Allied bombs rain down on Torino in the autumn of 1942, Stella Costa’s mother sends her to safety with distant relatives in a Tuscan villa. There, Stella finds her family tasked with a great responsibility: hiding nearly 300 priceless masterpieces from Florence, including Botticelli’s famous Primavera.

With the arrival of German troops imminent, Stella finds herself a stranger in her family’s villa and she struggles to understand why her aunt doesn’t like her. She knows it has something to do with her parents—and the fact that her father, who is currently fighting at the front, has been largely absent from her life.

When a wave of refugees seeks shelter in the villa, Stella befriends Sandro, an orphaned boy with remarkable artistic talent. Amid the growing threats, this unlikely found family takes refuge in the villa’s “treasure room,” where the paintings are hidden. There, Botticelli’s masterpiece and other works of art become a solace, an inspiration, and the glue that bonds Stella and Sandro as the dangers grow.

A troop of German soldiers requisitions the villa, beginning a dangerous German occupation that puts everyone to forced labor. Now, with the villa full of German soldiers, refugees, a secret guest, and hundreds of priceless treasures, no one knows who will emerge unscathed, and whether the paintings will be taken as spoils or become unintended casualties.